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一起神秘的死亡时间吸引了古生物学家凯丽（布里吉特·芳达 Bridget Fonda 饰）的注意，为了了解事件的真相，凯丽千里迢迢的赶到了案发地——一个古老的湖泊，而之后发生的事情更让凯丽确信，在这平静的湖面下，隐藏了什么可怕的东西。凯丽的猜测在小镇上引起了轰动，有些人信以为真，另一些人则不以为然。就在舆论的热度到达最顶峰之时，一个游客被不明物体拖入湖中的惨剧让所有人都意识到，事情严重了。 凯丽和管理员杰克（比尔·普尔曼 Bill Pullman 饰），警长汉克（布莱丹·格里森 Brendan Gleeson 饰）等人围绕着古湖展开了调查，令人震惊的是，在湖底，他们发现了一只异常凶猛巨大的史前巨鳄。
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Wealthy Rollo Treadway (Buster Keaton) suddenly decides to propose to his neighbor across the street, Betsy O'Brien (Kathryn McGuire), and sends his servant to book passage for a honeymoon sea cruise to Honolulu. When Betsy rejects his sudden offer however, he decides to go on the trip anyway, boarding without delay that night. Because the pier number is partially covered, he ends up on the wrong ship, the Navigator, which Betsy's rich father (Frederick Vroom) has just sold to a small country at war. Agents for the other small nation in the conflict decide to set the ship adrift that same night. When Betsy's father checks up on the ship, he is captured and tied up by the saboteurs. Betsy hears his cry for help and boards the ship to look for him, just before it is cut loose. The Navigator drifts out into the Pacific Ocean. The two unwitting passengers eventually find each other. At first, they have great difficulty looking after themselves, but adapt after a few weeks. At one point, they sight a navy ship and hoist a brightly-colored flag, not realizing it signals that the ship is under quarantine. As a result, the other vessel turns away. Finally, the ship grounds itself near an inhabited tropical island and springs a leak. While Rollo dons a deep sea diving suit and submerges to patch the hole, the black natives canoe out and take Betsy captive. When Rollo emerges from the ocean, the natives are scared off, enabling him to rescue Betsy and take her back to the ship. The natives return and try to board the ship. After a fierce struggle, Rollo and Betsy try to escape in a small dinghy. It starts to sink, and the natives swiftly overtake them in their canoes. Just when all seems lost, a navy submarine surfaces right underneath them and they are saved.